David Hembry

David Hembry

Assistant Professor
Biology Program
Department of Biology
ST 2230


  • PhD in 2012, University of California, Berkeley

Research Area

Evolution of species interactions, plant-insect mutualisms, coevolution, phylogenetics, macroevolution, evolution of symbiosis and parasitism, species interaction networks, leafflower-leafflower moth associations.

Current Teaching


  • General Biology II 
  • Evolution 
  • Animal Behavior
  • Vertebrate
  • Zoology

Select Publications

Chheang P*, Hembry DH, Yao G, Luo S-X. 2021. Diversity and species-specificity of brood pollination of leafflower trees (Phyllanthaceae: Glochidion) by leafflower moths (Lepidoptera: Epicephala) in tropical Southeast Asia (Cambodia). Plant Diversity doi.org/10.1016/j.pld.2021.07.001. [* co-supervised master’s student]

Hembry DH, Weber MG. 2020. Ecological interactions and macroevolution: a new field with old roots. Annual Review of Ecology, Evolution, and Systematics 51: 215-243.

Gloss AD, Nelson Dittrich AC, Lapoint RT, Goldman Huertas B, Verster KI, Pelaez JL, Nelson ADL, Aguilar J, Armstrong E, Charbonneau JLM, Groen SC, Hembry DH, Ochoa CJ, O’Connor TK, Prost S, Suzuki HC, Zaaijer S, Nabity PD, Whiteman NK. 2019. Evolution of herbivory remodels a Drosophila genome. Preprint available online at bioRxiv, doi.org/10.1101/767160.

de Aguiar MAM, Newman EA, Pires MM, Yeakel JD, Burkle L, Gravel D, Guimarães PR Jr, O’Donnell J, Poisot T, Fortin M-J, Hembry DH. 2019. Revealing biases in the sampling of ecological interaction networks. PeerJ 7: e7566.

Hembry DH, Raimundo RLG, Newman EA, Atkinson L*, Guo C*, Guimarães PR Jr, Gillespie RG. Does biological intimacy shape ecological network structure? A test using a brood pollination mutualism on continental and oceanic islands. Journal of Animal Ecology 87: 1160-1171. [* undergraduate co-author]

Luo S-X, Yao G, Wang Z, Zhang D, Hembry DH. 2017. A novel, enigmatic basal leafflower moth lineage pollinating a derived leafflower host illustrates the dynamics of host-shifts, partner replacement, and apparent co-adaptation in intimate mutualisms. The American Naturalist 189: 422-435.

Hembry DH, Althoff DM. 2016. Diversification and coevolution in brood pollination mutualisms: windows into the role of biotic interactions in generating biological diversity. American Journal of Botany 103: 1783-1792.

Hembry DH, Yoder JB, Goodman KR. 2014. Coevolution and the diversification of life. The American Naturalist 184: 425-438.

Raimundo RLG, Gibert JP, Hembry DH, Guimarães PR Jr. 2014. Conflicting selection in the course of adaptive diversification: the interplay between mutualism and intraspecific competition. The American Naturalist 183: 363-375.

Hembry DH, Kawakita A, Gurr NE, Schmaedick MA, Baldwin BG, Gillespie RG. 2013. Non-congruent colonizations and diversification in a coevolving pollination mutualism on oceanic islands. Proceedings of the Royal Society B 280: 20130361.

Hembry DH, Okamoto T, Gillespie RG. 2012. Repeated colonization of remote islands by specialized mutualists. Biology Letters 8: 258-261.

Last Updated: 11/17/2021